Department of Entomology


Joseph Morseucr entomology

Professor of Entomology

Location: Entomology 282
Tel: (951) 827-5814
E-mail: joseph.morse@ucr.edu

Research Specialization

My program covers both applied and fundamental research dealing with management of pests of subtropical crops, especially pests of citrus such as citrus thrips, Asian citrus psyllid, bean thrips, Fuller rose beetle, and various soft and armored scales, and pests of avocado such as avocado thrips, persea mite, and armored scales. I have an interest in integrated pest management, biological control, parasitoid biology and behavior, insectary rearing of natural enemies, the acute and sub-lethal impact of pesticides on both target pests and non-target organisms, modeling and computer simulation, pesticide resistance, and the management of invasive species. Over the past twenty years, my lab has contributed to the evolution of citrus and avocado pest management in California towards a more integrated approach emphasizing increased monitoring activity, use of economic thresholds and selective pesticides, conservation and augmentation of predators and parasitoids, and post-harvest disinfestation of fruit. I work closely with a number of citrus growers, pest control advisors, University extension agents, and researchers in other Departments on campus in order that our applied research efforts will have applicability to field pest management efforts. I have an interest in international agriculture and have been involved in citrus pest management projects and with researchers in Arizona, Florida, Hawaii, Texas, Argentina, Australia, Brazil, Chile, Cyprus, Egypt, Israel, Japan, Mexico, New Zealand, South Africa, and Spain, and in a 1996 FAO-sponsored analysis of citrus integrated pest management in 13 countries of the Near East. I have worked with the UC Statewide IPM Program (6 years as Associate Director), UC Center for Invasive Species Research (3 years as Director), and with UC's Division of Agriculture and Natural Resources (6 years as Program Leader for Agricultural Policy & Pest Management). I teach a course in the Natural History of Insects (Entm. 10) and the Evolution section of Introductory Evolution and Ecology (Biol. 5C) and enjoy graduate student instruction and interaction (7 M.S. and 10 Ph.D. students have been awarded their degree under my supervision).

Former Institution

Michigan State University


BS Electrical Engineering/Biology 1975
Cornell University
MS Systems Science 1977
Michigan State University
MS Entomology 1978
Michigan State University
PhD Entomology 1981
Michigan State University


2011 - Entomology Foundation Award for Excellence in Integrated Pest Management
2010 - Award of Honor, California Avocado Society
2006 - Art Schroeder Memorial Award, California Avocado Society
2006 - Fellow, American Association for the Advancement of Science
2005 - Award of Excellence, Citrus Research Board



Recent Journal Publications 

Harman, J.A., C.X. Mao, and J.G. Morse. 2007. Selection of Color of Sticky Trap for Monitoring Adult Bean Thrips, Caliothrips fasciatus (Thysanoptera: Thripidae). Pest Management Science 63(2): 210-216.

Byrne, F.J., N.C. Toscano, A.A. Urena, and J.G. Morse. 2007. Toxicity of Systemic Neonicotinoid Insecticides to Avocado Thrips, Scirtothrips perseae Nakahara (Thysanopera: Thripidae), in Nursery Avocado Trees. Pest Management Science 63(9): 860-866.

Kapranas, A., J.G. Morse, P. Pacheco, L.D. Forster, and R.F. Luck. 2007. Survey of Brown Soft Scale Coccus heseridum L. Parasitoids in Southern California Citrus. Biological Control 42(3): 288-299.

Prabhaker, N., J.G. Morse, S.J. Castle, S.E. Naranjo, T.J. Henneberry, and N.C. Toscano. 2007. Toxicity of Seven Foliar Insecticides to Four Parasitoids Attacking Citrus and Cotton Pests. Journal of Economic Entomology 100(4): 1053-1061.

Rill, S., E.E. Grafton-Cardwell, and J.G. Morse. 2007. Effects of Pyriproxyfen on California Red Scale (Hemiptera: Diaspididae) Development and Reproduction. Journal of Economic Entomology 100(4): 1435-1443.

Harman, J.A., C.X. Mao, L.J. Robinson, and J.G. Morse. 2007. Evaluation of Two Non-destructive Sampling Methods for Bean Thrips (Thysanoptera: Thripidae) Detection in Navel Oranges. Crop Protection 26(12): 1747-1754.

Triapitsyn, S.V., V.V. Berezovskiy, M.S. Hoddle, and J.G. Morse. 2007. A Review of the Neartic Species of Erythmelus (Hymenoptera: Mymaridae), With a Key and New Additions to the New World Fauna. Zootaxa 1641: 1-64.

Barry, J.B., S.B. Opp, J. Dragolovich, and J.G. Morse. 2007. Effect of Adult Diet on Longevity of Sterile Mediterranean Fruit Flies (Diptera: Tephritidae). Florida Entomologist 90(4): 650-655.

Krugner, R., M.W. Johnson, R.L. Groves, and J.G. Morse. 2008. Host Specificity of Anagrus epos (Hymenoptera: Mymaridae): A Potential Biological Control Agent of Homalodisca vitripennis (Hemiptera: Cicadellidae). BioControl 53(3): 439-449.  

Rill, S., E.E. Grafton-Cardwell, and J.G. Morse. 2008. Effects of Two Insect Growth Regulators and a Neonicotinoid on Various Life Stages of Aphytis melinus (Hymenoptera: Aphelinidae). BioControl 53(4): 579-587.

Kapranas, A., P. Pacheco, L.D. Forster, J.G. Morse, and R.F. Luck. 2008. Precise Sex Ratios Manifested by Several Encyrtid Parasitoids of Brown Soft Scale, Coccus heseridum L. (Coccidae: Hemiptera). Behavioral Ecology and Sociobiology 62(6): 901-912.

Krugner, R., M.W. Johnson, K.M. Daane, and J.G. Morse. 2008. Olfactory Responses of the Egg Parasitoid Gonatocerus ashmeadi Girault (Hymenoptera: Mymaridae) to Host Plants Infested by Homalodisca vitripennis (Germar) (Hemiptera: Cicadellidae). Biological Control 47(1): 8-15.

Humeres, E.C., J.G. Morse, R. Stouthamer, W. Roltsch, and M.S. Hoddle. 2009. Evaluation of Natural Enemies and Insecticides for Control of Pseudocysta perseae (Hemiptera: Tingidae) on Avocados in Southern California. Florida Entomologist 92(1): 35-42.

Haviland, D.R., S.M. Rill, and J.G. Morse. 2009. Southern Highbush Blueberries Are a New Host for Scirtothrips citri (Thysanoptera: Thripidae) in California. Florida Entomologist 92(1): 147-149.

Al-Wahaibi, A.K. and J.G. Morse. 2009. Temporal Distribution of Egg Hatch for Two Homalodisca spp. (Hemiptera: Cicadellidae) Under Constant Temperatures. Florida Entomologist. 92(1): 156-158.

Al-Wahaibi, A.K. and J.G. Morse. 2009. Egg Morphology and Stages of Embryonic Development of the Glassy-winged Sharpshooter, Homalodisca vitripennis (Germar) (Hemiptera: Cicacdellidae). Annals of the Entomological Society of America. 102(2): 241-248.

Al-Wahaibi, A.K. and J.G. Morse. 2009. The Egg Stage of Homalodisca vitripennis (Germar) and H. liturata Ball (Hemiptera: Cicadellidae): Ovometrics, Embryonic Development, and Non-parasitic Mortality. Annals of the Entomological Society of America. 102(2): 249-260.

Morse, J.G., P.F. Rugman-Jones, G.W. Watson, L.J. Robinson, J.L. Bi, and R. Stouthamer. 2009. High Levels of Armored Scales on Imported Avocados Raise Concerns Regarding USDA-APHIS’ Phytosanitary Risk Assessment. Journal of Economic Entomology 102(3): 855-867.

Krugner, R., R.L. Groves, M.W. Johnson, A.P. Flores, J.R. Hagler, and J.G. Morse. 2009. Seasonal Population Dynamics of Homalodisca vitripennis (Hemiptera: Cicadellidae) in Sweet Orange Trees Maintained Under Continuous Deficit Irrigation. Journal of Economic Entomology 102(3): 960-973.

Humeres, E., J.G. Morse, W. Roltsch, and M.S. Hoddle. 2009. Detection Surveys and Population Monitoring for Pseudacysta perseae on Avocados in Southern California. Florida Entomologist 92(2): 382-385.

Krugner, R., M.W. Johnson, D.J.W. Morgan, and J.G. Morse. 2009. Production of Anagrus epos Girault (Hymenoptera: Mymaridae) on Homalodisca vitripennis (Germar) (Hemiptera: Cicadellidae) Eggs. Biological Control 51(1): 122-129.

Rugman-Jones, P.F., J.C. Andersen, J.G. Morse, B.B. Normark, and R. Stouthamer. 2010. Molecular Phylogenetic Placement of the Recently Described Armored Scale Insect Abgrallaspis aguacatae and Several Congeners (Hemiptera: Diaspididae). Annals of the Entomological Society of America 103(1): 30-38.           

Al-Wahaibi, A.K. and J.G. Morse. 2010. Temporal Patterns in Homalodisca spp. (Hemiptera: Cicadellidae) Oviposition in Southern California Citrus and Jojoba. Environmental Entomology 39(1): 15-30.

Al-Wahaibi, A.K. and J.G. Morse. 2010. Biology of the Adult Stage of Ufens principalis (Hymenoptera: Trichogrammatidae), An Egg Parasitoid of Homalodisca spp. (Hemiptera: Cicadellidae) in Southern California. Annals of the Entomological Society of America 103(2): 236-246

Triapitsyn, S.V., P.F. Rugman-Jones, G. Jeong, J.G. Morse, and R. Stouthamer. 2010. Morphological and Molecular Differentiation of the Anagrus epos Species Complex (Hymenoptera: Mymaridae), Egg Parasitoids of Leafhoppers (Hemiptera: Cicadellidae) in North America. Zootaxa 2428: 1-21.   

Shi, X., J. Bi, J.G. Morse, N.C. Toscano, and D.A. Cooksey. 2010. Differential Expression of Genes of Xylella fastidiosa in Xylem Fluid of Citrus and Grapevine. FEMS Microbiology Letters 304(1): 82-88.        

Magsig-Castillo, J., J. G. Morse, G. P. Walker, J. L. Bi, P. F. Rugman-Jones, and R. Stouthamer. 2010. Phoretic Dispersal of Armored Scale Crawlers (Hemiptera: Diaspididae). Journal of Economic Entomology, 103(4): 1172-1179.

Al-Wahaibi, A.K. and J.G. Morse. 2010. Immature Development and Adult Eclosion of Ufens principalis (Hymenoptera: Trichogrammatidae), an Egg Parasitoid of Homalodisca spp. (Hemiptera: Cicadellidae) in Southern California. Bulletin of Entomological Research 100(4): 467-479.

Byrne, F. J., E. C. Humeres, A. A. Urena, M. S. Hoddle, and J. G. Morse. 2010. Field Evaluation of Systemic Imidacloprid for the Management of Avocado Thrips and Avocado Lace Bug in California Avocado Orchards. Pest Management Science 66(10): 1129-1136.

Kapranas, A., I.C.W. Hardy, J.G. Morse, and R.F. Luck. 2011. Parasitoid Developmental Mortality in the Field: Patterns, Causes and Consequences for Sex Ratio and Virginity. Journal of Animal Ecology 80(1): 192-203.

Rugman-Jones, P.F., L.D. Forster, E. Guerrieri, R.F. Luck, J.G. Morse, M.M. Monti, and R. Stouthamer. 2011. Taxon-Specific Multiplex-CR for Quick, Easy, and Accurate Identification of Encyrtid and Aphelinid Parasitoid Species Attacking Soft Scale Insects in California Citrus Groves. BioControl 56(2): 139-144.

Kapranas, A., D.L. Giudice, R.F. Luck, and J.G. Morse. 2011. Biology and Behavior of Metaphycus angustifrons Compere (Hymenoptera: Encyrtidae), a Newly Discovered Parasitoid of Soft Scale Insects (Hemiptera: Coccidae) in California. Biological Control 56(2): 139-144.

Prabhaker, N., S.J. Castle, S.E. Naranjo, N.C. Toscano, and J.G. Morse. 2011. Compatibility of Two Systemic Neonicotinoids, Imidacloprid and Thiamethoxam, with Various Natural Enemies of Agricultural Pests. Journal of Economic Entomology 104(3): 773-781.

Vasquez, C.J., R. Stouthamer, G. Jeong, and J.G. Morse. 2011. Discovery of a CI-inducing Wolbachia and its Associated Fitness Costs in the Biological Control Agent Aphytis melinus DeBach (Hymenoptera: Aphelinidae). Biological Control 58(3): 192-198

General Campus Information

University of California, Riverside
900 University Ave.
Riverside, CA 92521
Tel: (951) 827-1012

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Dept of Entomology Information

417 Entomology Bldg.

Fax: (951) 827-3086
Prospective Grad Students: (800) 735-0717
Grad Student Affairs